diff --git a/Lib/test/test_zipfile.py b/Lib/test/test_zipfile.py
index 29d98c8092..31c83987ab 100644
--- a/Lib/test/test_zipfile.py
+++ b/Lib/test/test_zipfile.py
@@ -1546,6 +1546,12 @@ class OtherTests(unittest.TestCase):
                 zinfo.flag_bits |= 0x08  # Include an extended local header.
                 orig_zip.writestr(zinfo, data)
 
+    def test_writestr_pathlike_issue40506(self):
+        with zipfile.ZipFile(TESTFN2, 'w') as orig_zip:
+            path = '/foo/bar.txt'
+            orig_zip.writestr(pathlib.PurePath(path), '1234')
+            self.assertEqual(orig_zip.open(path).read(4), b'1234')
+
     def test_close(self):
         """Check that the zipfile is closed after the 'with' block."""
         with zipfile.ZipFile(TESTFN2, "w") as zipfp:
diff --git a/Lib/zipfile.py b/Lib/zipfile.py
index 8903d6a42e..44b3ee8e63 100644
--- a/Lib/zipfile.py
+++ b/Lib/zipfile.py
@@ -341,6 +341,8 @@ class ZipInfo (object):
     )
 
     def __init__(self, filename="NoName", date_time=(1980,1,1,0,0,0)):
+        if isinstance(filename, os.PathLike):
+            filename = os.fspath(filename)
         self.orig_filename = filename   # Original file name in archive
 
         # Terminate the file name at the first null byte.  Null bytes in file